Howarth, F.G. (1987a). The evolution of non-relictual tropical troglobites. International Journal of Speleology 16: 1–16.
| Taxon | |
|---|---|
| Tyrannochthonius pupukeanus Muchmore, 1983 (Pseudoscorpiones: Chthoniidae) | 3, Tyrannochthonius pupukeanus |
| Vulcanochthonius howarthi (Muchmore, 1979) (Pseudoscorpiones: Chthoniidae) | 3, Tyrannochthonius howarthi |